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Elite Member
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Join Date: Mar 2016
Location: Pasadena, CA
Posts: 5,409
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Elite Member
Join Date: Mar 2016
Location: Pasadena, CA
Posts: 5,409
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The ECU controls transmission shifting from sensors exclusively inside the transmission. It still expects to see a speedometer signal separately from the gauge cluster (not used for shift logic through) - and will throw a code if that signal is missing for a while, but not immediately.
Look for a '99/'00 cluster. Since you have an SR5, you can get the cluster from any V6 auto 2WD or j-shift 4WD.
If you do find one, the first test would just be to run the new cluster to see if that fixes the problem. The next step would be figuring out if you can make a frankenstein cluster from the two that will work and keep your odo correct.
-Charlie
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'99 4Runner SR5 Auto - 4WD swapped
'89 Camry Alltrac LE 3S-GTE 5spd
'17 Chevy Volt Premier
'16 Honda Odyssey Elite
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