I have been researching tires for 3-4 months and my driving habits and needs are pretty much identical to yours. 90-95% on road with occasional forest service roads and some winter driving in the mountains for backpacking, fishing etc. I had KO2’s on my f150 and they got scary in the rain as the miles started to add up. Unless sidewall protection is crucial they are overkill for a 4runner. Especially if wet weather traction is key and I see tons of rain and wet roads in upstate SC and the N.C. mountains. I ran p metric michelins and Yokohama geo landers for many years on my Gen 3 and never had a single issue.

The Falken WP is probably going to be my next purchase. I like the aggressive look and they check all the boxes. They are 7lbs heavier per tire which is the only drawback vs the stock Dunlops. Other tires I have considered are the cooper at3 and the Toyo OC at2. The coopers just don’t appeal to me as much visually. The Toyo’s get mixed reviews but in P or SL load they do weight 41 lbs so some benefit there from a mpg perspective.

I plan to take the mileage hit and go with the Falkens. Hope this helps you out.