Well I understand the Speedo/ABS relation, however, ignoring the speedo for a minute and referring to the GPS speed I'm getting the same value as before swapping over the third member. That tells me the tires are spinning the 4Runner faster forward, with my original 3rd member and the one I swapped in giving the same result.
OK, so I'm going to just count the teeth of my original 3rd member to find out once and for all what it actually is, and going with the logic that the 3rd member I swapped in is identical. I'll report back once I've counted!