Sounds similar to what I was hearing. On mine it was the lower control arm bolts. The alignment technician left them a little more than snug. I torqued them and the sound is gone. I'm probably out of alignment again, though.
We did a similar trip this summer. Rhode Island to Cape Breton Island to Prince Edward Island and back. We had only bought the 4runner a couple of weeks before this trip. I had to haul a lot of extra stuff because we had 2 kids and a dog taking up the cargo space. Kind of missed the Tacoma...
I can't speak about the KO2, but I have owned 4 trucks with KOs. I have also run Toyo open country at2s, cooper discoverer at3s and several other popular tires. I am currently on Falken wildpeak at3w. I was unsure about my decision at the time I purchased them, but I have been impressed...