The nav head unit is only on or off. The base radio has like 3 or 4 settings: high, med, low or off. I don't notice it raising the volume or anything when it's on and I still fiddle with the volume from time to time depending on city or freeway driving, maybe my MP3s' volumes aren't normalized. However, since the 4R is much quieter than most vehicles, city volume levels can be heard quite well on the freeway.
Play around with it, you may like the feature (use to be only on Lexus vehicles) or you might just find it like me--makes no significant difference on or off. I just leave it on.