My son went from 3rd gen to FJ. I have 5th gen, and previously 4rth gen. He's driven them all. He loves them all, but every time he gives me my keys back he tells me how much he misses 4runner. He loved windows down, sunroof open, and rear window open running. It apparently is a big thing for him.
Personally, I hate the lack of visibility in his FJ. Particularly where it pertains to having to lean forward to see the stop lights.
Yes I am more prone to high center, but I've made it out of the mud where he couldn't. Wider wheel base?