so you're saying right and left wheels are locked together with center diff lock?
No.
Rear diff lock and center diff lock are apples and oranges.
Here's why:
First off, I'm sure you understand the purpose of the differentials in your axles, right? When you go around, let's say, a left hand turn, your outer (right) wheel has to go faster and farther, while your inside (left) wheel goes slower and a shorter distance. The differential is what allows the outer wheel to go faster and the inner wheel to go slower.
If you didn't have a differential there at all, you'd have a locked axle - the left and right would get the same amount of power and would always turn at the same speed.
The problem with open differentials is that if you lift one wheel off the ground, it will take all the power, and it will spin in the air while the tire with traction remains motionless on the ground.
What a rear locker does, then, is that it temporarily disables the rear differential and allows both wheels to turn at the same speed all the time. Obviously, this is not something you want to do on the streets as it could cause damage to your tires and drivetrain, but off road when you are bouncing through the rocks it can be a lifesaver.
The center differential is different.
On a "conventional" 4wd vehicle, you cannot run the vehicle in 4wd on dry, hard surfaced roads. The reason for this is that the transfer case splits power 50/50 to the front and rear driveshafts. The problem with that is that on most vehicles, when turning a curve, the front axle actually travels a bit farther than the rear axle. So, with 50% of the power going to the front axle and 50 to the rear, both axles (and the rest of the drivetrain) get stressed out because the front axle has too little torque and the rear axle has too much. That is why conventional 4wd vehicles all have stickers or plates that warn you not to drive in 4wd on a dry, hard surfaced road.
Now, on a wet, slippery road, or off-road, this isn't a concern because the slipping caused by the lack of traction will relieve the drivetrain of any stress. Off-road, if you lift your wheels (pretty common off road) this will also "unload" the stressed out drivetrain, which is why you can run in 4wd all day if you are 'wheeling.
Years ago, manufacturers saw the desirability of running in 4wd on the streets, so they came up with a system where they put a differential (like the one you have in your front and rear axles) into the transfer case. The center differential allows the front and rear driveshafts to turn at slightly different speeds to allow for the longer path of the front axle. These vehicles are full-time 4wd or are sometimes called AWD.
The problem with the center diff is that, like the open diff on the axle, it wants to send all the power to the driveshaft that is spinning the most, so if you lift one front tire off the ground and it takes all the power from the front axle, the center differential will want to take all the power, too, and leave the rear driveshaft (and rear axle) with no power. So, almost every manufacturer who makes a full-time 4wd vehicle equips it with a center differential lock. The center diff lock disables the center differential and the vehicle, at that point, acts just like a conventional 4wd vehicle with 50% of the power going to the front driveshaft and 50 to the rear. Of course, this is for off-road/low traction surfaces only.
On the 4runners, All 4runner through 1998 had conventional 4wd. No center diff lock.
The rear E-locker became available in 1996 with the 3rd gen 4runner. The E-locker is on most Limiteds and some SR5s (my SR5 has one.) E locker was available through the 2000 model year.
From 1999 through 2000, standard models and SR5s had conventional 4wd with no center diff lock. Limiteds (most of them anyway) had
both the rear E-locker
and the center differential (which is one reason why the 99 - 2000 Limiteds are among the most desirable of 3rd gen 4runners.)
Starting in 2001,
all 4runners got the center differential, but the E-locker was no longer an option. They also got a VSC (Vehicle Stability Control) system.
Put more simply: The center differential gives more
on-road capability (by allowing you to drive your vehicle in 4wd on the streets) while the rear diff E-locker gives better
off-road performance by allowing you to lock your rear differential when in 4wd-lo range.