This takes me back to a question about the center diff lock which I dont think I posed correctly. I understand how a rear diff lock works.
My question is, what is the benefit of a rear locker in addition to the center lock on an 09 Trail?
I ask this because in 4HI or 4LO in my 4th gen, if you take grip away for 2 diagonally opposed wheels, or suspend them completely, they will spin. With the center diff locked, they will not. At least I have never experienced a situation where there is any slip with the center diff locked. Its easy enough to understand a schematic on how rear lock works, but I cant get my mind around how the center lock and front and rear axles/driveshafts are connected. (I am not asking you to try and explain!, If you have a link, pls send).
But what is a situation that a rear diff lock would help, where a center lock would not?
Center diff lock = locks the front and rear axle together; thus, torque flows to the front and rear axle equally. (Coincidentally, it also shuts off VSC.)
On the 4th gen, you have a Torsen center differential. It allows for torque to go to the front and rear differential on dry land. However, how much of the torque to each axle varies upon road conditions. Sometimes, it will send 70 percent of torque to the rear axle (and 30% to the front)...such as when you turn on a curve in the road. Other times, it will send 60% of torque to the front axle (and 40% to the rear)...such as when one of the tire in the rear axle is slipping. Yet, other times, it will send 50%/50% to each axle when you're driving straight.
However, in off-road conditions, you do not want the computer to think too much...you do not want it to shift power to either axle when you don't need to. In addition, the Torsen center diff has a nasty habit of sending ALL power to the wrong axle IF both tires on one axle has lost complete traction. So, if your rear axle is sunken in deep mud and has no traction whatsoever, then the Torsen will INcorrectly send all power to the rear axle...thus, you're stuck. This is why LOCKING the Torsen off-road is a good thing...to prevent such issue and to take the ECU (computer) out of overthinking itself. (As noted above, locking the center diff will also turn OFF stability control, which tends to de-throttle your engine when it encounters slippage.)
Back to your original question. I think that it is coincidence that you see less slippage with the center diff locked. Center diff only controls power flow to the front and rear axle. The power flow laterally (left and right) on EACH axle is controlled by the traction control system (ATRAC). Therefore, in your diagonally suspended 4runner, it is the ATRAC that makes the most difference in slowing down the wheels that are in the air and shifting power to the wheels that are on the ground. The locking center diff is just there to guarantee that each axle will have 50% of engine torque for ATRAC to shift power left and right. BTW, for ATRAC to work its magic, there MUST be some slippage...because the wheel sensors (via ABS logic) MUST sense wheel speed differences between the wheels on the SAME axle for it to engage.
In other words, if your 4runner did NOT come with ATRAC and has totally open differentials front and rear, then you would be stuck in the diagonal situation you described, regardless of whether your center diff was locked or not.
A rear locker is good off-road because, in your diagonal situation, the rear axle tires do NOT have to wait for slippage for both tires to get power. Lets concentrate only on the rear axle. With ATRAC controlling the rear axle, there must be slippage (from the tire suspended in air) before ATRAC will slow that tire down. This will then allow SOME torque to flow to the other tire on the rear axle that is touching the ground. So, there will be a moment of slippage before you see any power flowing to the other tire. A rear locker locks the two rear wheels together. Therefore, both tires are ALWAYS getting 50% of torque that is being sent to the rear axle via the center diff. There is no waiting for one tire to slip before the other gets power...both tires ALWAYS get power. Thus, in your diagonal situation, there won't be any hesitation (or waiting) before your 4runner will move forward. This is why a locker is superior to ATRAC under most off-road conditions.
In summary, a center diff lock does NOTHING to the torque flow between your left and right tires. Its only job is to make sure that power flows to both front and rear axles equally (north and south)...and that VSC (stability control) does not take power away from you. ATRAC is what controls the power flow between your left and right tires on each axle via pulsating the brakes on the slipping wheel (east and west)...it requires tire slippage before it will direct power to the proper tire. A locker does the same thing as ATRAC but it does not require tire slippage before it engages...it is always sending power to all tires on the same axle that is being locked down.
Hope this helps in some way...sorry if i went a bit off-topic.