Off the top of my head, heated steering wheel, HUD, memory seats, MUCH better resolution cameras, camera rear view mirror, panoroof.
I think it just comes down to a person's priorities, their use case, and where they see value. To me charging $55k for a 4-runner that has less tech in it than a $40k Rav4 makes a statement. For example I used to make fun of heated steering wheels before I had one for a winter. That said I could care less about parking sensors and radar cruise, trade them both for a HUD and heated steering wheel in a heartbeat.
If my $58k 1/2 ton pickup can find a way to get 420hp/460ft/lbs of torque (at nearly the same MPG), HUD, tow 9000 lbs, heated steering wheel, heated/ventilated memory seats, camera rear view mirror, nice resolution 360 cameras, memory seats at almost the same price, I don't see much value in the current 4-runner at it's price. I can respect minimalist for reliability sake, but not combined with a premium price tag.
Unfortunately we're more likely to see more and more cars neutered when it comes to off-road and features. The 2023 Jeep lineup is a perfect example, they neutered the Grand Cherokee line unless you buy the hybrid trims now. The only trailhawk version is now the 4xe, and you can't get the off-road package on any Grand Cherokee now unless it's a 4xe. The only non-hybrid Grand Cherokee that has the eLSD now is the Summit Reserve and it comes with 21" rims.
I love the looks and reliability of the Toyota, but when you compare the $55k TRD Pro to something like the Grand Cherokee Trailhawk for another $5k the Trailhawk gives you the same.....I'll even say significantly more off-road prowess in stock form (11.3" ground clearance -vs- 9.6", 35 -vs- 33 approach angle, 25 vs 20 deg breakover, 30 deg vs 26 deg departure angle, similar if not better RTI (depending whose #'s you use)). Plus you have 6200lbs vs 5000lbs of towing, or 7200lbs if you get the v8 and all the creature comforts inside. Now of course you can't mod up a Grand Cherokee like you can a 4-runner, but for $60k I'd wager it will embarrass stock TRD Pro's off road. The only thing I can think of that's not in the Trailhawk is you can't get a HUD in it. Interestingly they also have basically the same tire size, except the 4-runner is on 17" rims and the Trailhawk is 18". On road and comfort/convenience/tech wise it's not even a contest, the Trailhawk is vastly superior. The only place the 4-runner does better is of course rugged looks, we can all agree it's probably more reliable, mod capability you can turn a 4-runner into a monster, and it's got about 10% more cargo room. The trailhawk will even give you stuff most of us don't care or need.......night vision, memory steering column, it also has more HP and Torque even with the v6 and better fuel economy.
For me the real problem with the 4-runner is I'm not going to turn it into a modded monster, if I was going to do that I'd probably just mod out a jeep. So I'm not going to do much to it beyond stock, and it's going to be a daily driver, so if it's a worse vehicle on road, less comfortable, doesn't drive as nice, has less tech options, less convenience. The only thing that would get me to go that way is if it's super reliable (check!) and it had a significant price break compared to vehicles with all those things, and it just doesn't. Even if you say went to the TRD Off Road Premium, and wanted KDSS to improve the on-road manners you are looking at $50k. Now if that was closer to $45k then you've got a pretty good price gap to something like the Trailhawk.
The other advantage of course is resale, in 5 years the 4-runner is going to hold it's value MUCH better, Jeeps are one of the worst at holding value. However I've always said if you buy a vehicle for it's resale, you probably bought the wrong vehicle, but I've been driving for 34 years and have only owned 4 vehicles, so resale for flipping purposes isn't a factor.