Between the 4Runner and Land Cruiser, price is certainly one thing and MPG is not far off in the same vein. Being able to get close to 20mpg on the highway with the 4Runner really ads up when you consider 15mpg on a Land Cruiser.
With gas less than $2/gallon, I don't care about the difference right now. Of course, gas probably won't stay this cheap.
I commute in mostly stop and go traffic into Cambridge every week day. My 4Runner got about 15.5 mpg in that commute and about 17 mpg on the highway.
My Land Cruiser is averaging 14 mpg on that same commute, so it is worse than my old 4Runner, but not by a lot. I have only done one "highway" trip with my Land Cruiser, but it involved a lot of rural two-lane highways in Vermont (lots of hills), and on that trip I got 17 mpg. So I think on an interstate I could probably get 18 mpg.
Real world combined fuel economy for the 5th Gen 4Runner seems to be around 18 mpg. I doubt I would see that on my stop-and-go commute, but let's just assume that I would and look at the numbers.
I drive about 15k miles per year. If I get 14 mpg combined, then at $2/gallon my fuel cost is $2143/year.
If I was driving a 4Runner and getting 18 mpg combined, then at $2/gallon my fuel cost is $1667/year, for savings of $476.
If gas goes back up to $4/gallon then those costs are double.
Btw, the Fast Lane Truck guys were able to get 20 mpg out of the 2016 Land Cruiser on the highway at 70 mph. That number was determined by hand (miles driven divided by gas pumped). That is pretty darn good for a truck this big and heavy.